MR/P024335/1Medical Research Councilhttp://dx.doi.org/10.13039/501100000265MR/P024335/1Arts and Humanities Research Councilhttp://dx.doi.org/10.13039/501100000267MR/P024335/1Biotechnology and Biological Sciences Research Councilhttp://dx.doi.org/10.13039/501100000268MR/P024335/1Engineering and Physical Sciences Research Councilhttp://dx.doi.org/10.13039/501100000266MR/P024335/1Natural Environment Research Councilhttp://dx.doi.org/10.13039/501100000270NE/R000131/1Natural Environment Research Councilhttp://dx.doi.org/10.13039/501100000270